What are the other alien skulls in the Predator trophy case?

A well-known easter egg from Predator 2 is the presence of an Alien skull in the trophy case on board the Predator ship. Aside from a couple of human skulls, there are 4 others:

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And here's a close-up of the two skulls abutting the Alien skull:

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Can any of these skulls be identified as either homages to other sci-fi creatures (like the Alien skull) or belonging to creatures appearing in any of the subsequent Predator comics, books, or other movies? Or are they simply props appearing in just this one movie?

Answer

They are the unidentified species which are throughout the Aliens / Predator universe.

The skulls were likely created in order to demonstrate that the Predators are not just predators of men, but of the whole galaxy. They travel the stars looking for the "great hunt."

In the second photo, the left skull has some passing likeness to a River Ghost:

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