a song of ice and fire - What was the original cause of enmity between Stark and Lannister?

In A Game of Thrones, chapter 4 (the first Eddard chapter), we hear Eddard and King Robert talking about sending Lysa Arryn's son, Robert Arryn to be a ward of Tywin Lannister. Eddard is relieved that Lysa had declined:

Ned would sooner entrust the child to a pit viper than Lord Tywin, but he left his doubts unspoken. Some old wounds never truly heal, and bleed again at the slightest word.

- AGOT Chapter 4

What "old wounds" is Ned thinking about here? In the next Eddard chapter we learn that Ned distrusts Tywin for ordering the murder of Rhaegar Targaryen's children. But the way Ned thinks of Tywin in the quote above makes me think the wounds are deeper and more personal than that.

There is obviously enmity between the Starks and Lannisters after the events of A Game of Thrones but what happened to cause enmity before AGOT?

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