In The Matrix Revolutions, when Morpheus, Trinity and Seraph enter the Merovingian's club, one of the guards calls Seraph "Wingless". Where does this name come from?
Answer
As his name suggests, Seraph is believed to be a Seraphim (a type of program that performed the function of Agents in the Paradise Matrix). Seraph used to work for the Merovingian but was exiled along with the Merovingian and his other henchmen.1 Eventually he came to protect the Oracle, thus betraying the Merovingian2 and losing his exile status (since he had a purpose again).
At some point Seraph lost his wings, possibly from an attack by the Merovingian similar to the latter's attack on the Oracle which forced her to change her shell. He can no longer fly, in contrast to the guards inside Club Hel who work for the Merovingian and can defy gravity -- they are fallen angels (in Hell no less!) who still have their "wings" (although we can't see them).
(You can learn more about the history of Seraph, the Oracle, and the Merovingian here.)
1We know this because Smith tells Seraph in The Matrix Revolutions that "chasing you was like chasing a ghost", and it is part of Agents' responsibility to apprehend exile programs.
2 This is why the Merovingian calls him the "prodigal son" and "Judas" in Club Hel.
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