I don't remember in which movie it was mentioned for the first time.
It was some movie with John Malkovich if I remember well.
Answer
"The Hitchhiker's Guide to the Galaxy" was a 2005 movie where John Malkovich played "Humma Kavula" (Zaphod Beeblebrox's competitor for President of the Galaxy post).
That movie was based on Douglas Adams' "The Hitchhiker's Guide to the Galaxy" book, which famously originated "42" as the "Ultimate Answer to the Ultimate Question of Life, The Universe, and Everything".
The details on the Ultimate question are covered on Wiki
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