In the 2007 movie, Optimus explains that the Autobots learned to speak English from the world wide web. One could reasonably assume that the Decepticons did as well.
But Megatron wakes up after being trapped in stasis within the ice and says, "I am Megatron!" and then goes outside and speaks with Starscream in English.
There is at least one scene where the Decepticons (Starscream, I believe) speak in Cybertronian and they display English subtitles, but they don't do that with Megatron.
Megatron speaks to Sam in English as well, so it's not just a matter of languages being translated for the ease of the viewer following along.
Answer
This is simple supposition, but I would imagine that the language could be communicated fairly easily from Starscream(or another Decepticon) to Megatron. These are advanced machines, it isn't like he has to put his spectacles on and sit down with a copy of English for Dummies, he should be able to just download the language files from another.
This doesn't necessarily explain why he immediately does this and speaks English to Starscream, but that particular instance could be explained by your last statement (ease of use for the viewer following along).
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